The four-inch-tall Clock family secretly share a house with the normal-sized Lender family, "borrowing" such items as thread, safety pins, batteries and scraps of food. However, their peaceful co-existence is disturbed when evil lawyer Ocious P. Potter steals the will granting title to the house, which he plans to demolish in order to build apartments. The Lenders are forced to move, and the Clocks face the risk of being exposed to the normal-sized world.

The Borrowers are small, 15cm high humans who live in the English hinterland. They live out their lives in mouse-hole sized nooks in human homes, and survive by 'borrowing' all they need from the house and its inhabitants. This series follows young girl Arriety, and her parents Pod and Homily, as they are displaced from their home and try to find a new home, with the help of a human boy, George.