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Autobots, Decepticons, Predaking and Predacons form an unlikely alliance to battle a resurrected Unicron.

Aidan, an awkward fanboy, finally leaves his mother’s basement and goes to meet Sierra, a vlogger fresh off a break up who needs help filming a new video series at Comic-Con. They struggle to navigate the newly devirtualized relationship, and just as they’re starting to figure it out, disaster strikes. Further alienated and unable to cope, Aidan descends into nightmares, becoming a monstrous toy as he dawns Sierra's internet identity and forges on to Comic-Con to complete the video series.

Sam Witwicky leaves the Autobots behind for a normal life. But when his mind is filled with cryptic symbols, the Decepticons target him and he is dragged back into the Transformers' war.

The Autobots continue to work for NEST, now no longer in secret. But after discovering a strange artifact during a mission in Chernobyl, it becomes apparent to Optimus Prime that the United States government has been less than forthright with them.

As humanity picks up the pieces after the battle of Chicago, a shadowy group reveals itself in an attempt to control the direction of history…while an ancient, powerful new menace sets Earth in its crosshairs. With help from Cade Yeager, Optimus Prime and the Autobots rise to meet their most fearsome challenge yet.

Humans and Transformers are at war. Optimus Prime is gone. The key to saving our future lies buried in the secrets of the past, in the hidden history of Transformers on Earth. Saving our world falls upon the shoulders of an unlikely alliance: Cade Yeager; Bumblebee; an English Lord; and an Oxford Professor.